Yes, the physiology and biomechanics of treadmill running are a little different. But how you feel about it is probably more important.
It’s probably true that people run slightly differently on treadmills: when you look in detail at the biomechanics, you find subtle differences in things like knee angles and peak ground forces, but the overall pattern (as one such study concluded) is that the two movements are close enough that you don’t have to worry about the differences—as long as the belt is properly calibrated and not too soft.
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